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arachidonic acid ara aa polyunsaturated omega-6 fatty acid omega eicosatetraenoic arachidonate cupuacu butter peanut oil double bond phospholipid phosphatidylethanolamine phosphatidylcholine phosphatidylinositide membrane cellular signaling vasodilator essential acids vitamin biosynthesis alzheimer's disease supplement dietary nutrient pc lecithin sphingomyelin choline arachidonic lipid phosphocholine cholesterol fatty ceramide lysophosphatidylcholine bilayer surfactant glycerophospholipid phosphatidylserine lipophilic 3d