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virodhamine o-arachidonoyl ethanolamine o-aea endocannabinoid eicosanoid arachidonic acid endocannabinoid system antagonist neurotransmitter receptor agonist neuromodulatory nervous brain stress appetite memory signaling neuroscience cb2 cb1 cannabinoid activity receptors physiological cell membranes ecs neural addictive-like behavior pain modulation learning emotion motivation motor coordination molecule molecular formula skeletal structural structure geometry ball stick 3d bioactive cell biochemistry biomolecule