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androstenol steroidal pheromone neurosteroid kayromon allomone musk-like odor black truffles perfume effects foods smell fragrance attraction benefits positional isomer biosynthesis axillary sweat blood-brain-barrier central nervous system hormone hormonal function action endocrine prohormone analogue molecule molecular formula skeletal structural structure metabolite metabolism mechanism biosynthetic biogenesis substrate precursor synthesis derivative bioprecursor metabolic endogenous 3d