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acetyl-coa a-ac-coa acetyl coenzyme biochemical reaction protein metabolism citric acid cycle krebs oxidized coash mercaptoethylamine group pantothenic acid high energy glycolysis carbohydrates atp gtp synthesis extramitochondrial enzyme atp citrate lyase intramitochondrial pyruvate dehydrogenase complex metabolite biosynthesis mechanism pathway path biosynthetic intermediate biogenesis substrate precursor derivative bioprecursor metabolic enzyme endogenous steroid acetylcholine melatonin synthesis acetylation allosteric regulator 3d