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acalyptratae animal animals aquatic arthropod bug couple coupling cyclorrhapha detail diptera ephydridae ephydroidea fauna fly freshwater gerridae gerrinae gerrini gerris gerroidea hemiptera hemipteran heteroptera insect invertebrate lake liquid macro natural nature pest pond predator reproduction river schizophora sex skater spider strider water waterbug waterstrider wet wild wildlife yellow typical genus neogerris