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marsh samphire sea beans salicornia herbacea vegetative settlement dry land drought spell salttolerant plant salt immunity puffed solonchak extreme salinization saline soils alkaline earth solonchaklike soil alcali alkali spot crust flat wet sulfide silt deposits sarzasan halophylic vegetation halophytes semidesert halophytic drimophilus halophile petrosimonia suaeda solecornia prairie soda xerophyte xerophite succulent saltwort quickly colonized europaea spring flood drought salt-tolerant