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nyala tragelaphus angasii spiral-horned antelope native southern africa species family bovidae genus considered nyala antelope tragelaphus angasii buck lowland horn spiral animal mammal nature wildlife africa female natural bovidae wild south african south herbivore kruger stripes white outdoor park safari savanna wilderness botswana conservation reserve horned kenya brown national antilopes etosha forest namibia artiodactyla animals hoofed zimbabwe background grass