The Chestnut-mandibled Toucan, or Swainson's Toucan (Ramphastos swainsonii). Sometimes individual fruit trees are defended by a mated toucan pair from other toucans or from other frugivorous birds - defended by threat displays and even, against other toucans, by bill clashes. Chestnut Mandibled toucans, the largest in Costa Rica, may parasitize the slightly smaller Keel-Billed Toucan. The larger
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