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asteraceae compositae serratula tinctoria anthesis capitulum corymbose disc floret flower gynoecium head imbricate inflorescence involucral bract involucre paniculate phyllary stigma style subtending synflorescence tubular sawwort dyer 039 plumeless saw-wort tinctoria closeup virtual herbarium belarus flora: focus-stacked closeups wild cultivated plants including flowers leaves fruits intended illustrations botanical manuals