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bird rufousbacked wren costa rica campylorhynchus capistratus nature ocotal coco tropical fauna avian colorful vibrant feathered perched plumage rainforest biodiversity exotic wild avifauna central america beauty endemic songbird habitat small natural rican feathers diversity watching ecosystem ecotourism delicate stunning ecological species wilderness ornithology birding birdwatching animal wildlife rufous-backed capistratus beach family troglodytidae