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saxifrage silene acaulis diapensia pincushion plant pillow shaped vegetation circumpolar arctic alpine lapponica perennial plants herbaceous blooming stems environment lapland tundra mountain fjelds cold desert bald landscape ecology erica fenland floral flower foliose lichen grass green grower habitat herb lichens loiseleuria marchantiophyta north northpolar association polar polarizer prairie scablike lapponica start remains last year