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39aph39 africa african arid bill bird black bucerotidae casque claws curved decken deckeni der dimorphism droppings east eggs entrance ethiopia excreta family feathers feed feeds female flight flocks forests fruit grasslands ground habitats hole hornbill incubation insects long male mud nature nest omnivorous plaster pulp red rift rodents s savannahs scrub sealed seeds sexual slit small tail tanzania thorn tockus tree tropical upperparts vertical von white wings