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iron distribution transferrin ferritin hemoglobin tissues organs liver bone marrow spleen macrophages erythropoiesis cellular uptake transport systemic circulation storage heme synthesis red blood cells hepcidin homeostasis metabolism ferroportin duodenum absorption recycling ironrich foods anemia deficiency receptors intracellular ironexporting proteins regulatory ironbinding capacity serum nonheme dietary intake tissuespecific regulation utilization breakdown stores turnover balance release sequestration export transporters involves metabolic processes body organs ilustration