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animal insect invertebrate wildlife nature leaf flower arthropod branch plant beetle food tree pest produce horn bangbung oryctes rhinocerus type rhinoceros dynastinae widespread southeast asia eastward island guinea northward formosa adults black dark brown body length horned head horns male beetles longer curved female bulges larvae big milk gray-white covered hairs small reddish-brown feeds pith material leaves attach tips absorb fluids damage young tissue common types palm trees palmae stems host plants