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hamedafne ericaceae russia flowering heather dicotyledonous plants hamedafna chamaedaphne karelia onezhye ecosystem flora leatherleaf bereinis chamedafne vaivero torfgranke swamp forest waterlogging arctic north evergreen bush orzega mycotroph glycoside andromedotoxin neurotoxin acetylandromedol rhodotoxin polyhydroxylated cyclic diterpene