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guadiana spanish pond turtle shell natural light closeup extremadura saharan mauremys leprosa long format panoramic reptilie testudines iberian mediterranean geoemydidae hard empty animal semiaquatic fauna freshwater carnivorous brown vertebral keel patent worn appearance remains omnivorous asymptomatic carriers salmonella bacteria shield dorsal parts dermal bone enclosure protection identification tool keratinous scutes carapace structural support back convex ossified ribs fused ossification skin dry horny plates structure terrestrial shell light