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palythoa zoanthid zoantharia zoanthidea zoanthinaria anthozoa zoanthus hexacorallia cnidarian coralimorph coelenterates coenenchyma zooxanthellae palytoxin toxic poison tentacle stalk capitulum detritus zooplankton intertidal subtidal current aquatic biodiversity fauna ecosystem environment underwater captive wildlife aggregation lateral saltwater aquascape reef tank aquaculture tropical macro encrusting controlled conditions shallow dof space blur background glass refraction propagation demanding