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abdomen altruism animal ant antenna antennae antics ants arthropod background biodiversity biome camouflage colony communication defense ecosystem egg entomologist entomology eusocial exoskeleton foraging formicidae habitat hibernation hill hymenoptera insect invasive larva macro mandible mound myrmecology myrmicine nature nest pheromone predator prey queen small social soldier species symbiosis thorax tree worker