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grant's zebra equus quagga boehmi plains serengeti-mara ecosystem vertically striped stripe horizontal back diagonally rump shadow family group stallion wild equidae equids mammal ungulate pattern black white brown herbivore grass migration harem herd water animal forage savannah poaching lion predator unique strong eyesight hearing savanna grassland shrubland berries plant woodland habitat coarse vegetation central africa northern smallest subspecies social zambia congo tanzania kenya ethiopia somalia