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exosome intercellular communication coagulation vesicle extracellular mhc2 pdl1 egfr integrin tetraspanin mrna galectin chaperones ubiquitin lipid bilayer membrane biological protein molecules exosomes cross showing vesicles involved physiological processes like waste management bound proteins mhc-ii pd-l1 integrins tetraspanins carry signaling vegf galactin red