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curly lobe helvella crispa mushroom species genus helvelaceae family helvellaceae lectotype fruiting body apothecia saddleshaped lobed spores forest foliage autumn saprotroph mycorrhizaforming mushrooms eukaryotes ascomycete pecicomycete pecicaceae gelvellids serbia balkans bosnia and herzegovina guevo september europe ecosystem ecology bio eukaryotic organisms signs of plants animals mycology botany biology dry decomposer biosphere symbiotic mutualistic heterotroph mycelium thallus substrate rhizomycelium