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actiniidae amphiprion aquaculture aquascape background biocenosis biodiversity bulbous clarkii clownfish cnidaria commensal community conditions controlled coquina coral coralimorph cultivation current dark decorative depth dominance ecology ecosystem entacmaea environment exotic farm fauna field focus free hermaphrodite hierarchy indopacific mutualism mysterious nature ocean photosynthetic plankton poison pomacentridae protection quadricolor reef salt saltwater selective sequential shallow space symbiont symbiotic tank tip tropical understanding underwater venomous wildlife yellowtail