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gfap glia immunohistochemical ihq bergmann central nervous system cerebellar cortex cerebellum cns light microscope micrograph microscopic microscopy tissue neuroanatomy neurohistology neuroscience histology histological immunohistochemistry cortex immunostained for glial fibrillary acidic protein marker astrocytes that are located granular layer molecular layer parallel radial processes reach pial surface belong golgi epithelial cells