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adaptation aiding anatomy annulus arm armor bead behavior biology bodies buoyancy camouflage carp carpio closeup covering ctenoid cycloid cyprinus defense dermis diversity ear ecology epidermis estimation eye factors finger fish fisheries food functions gemstone grow growth hand ichthyology important including interactions jewellery leg limb lip management mineral nail natural nature organ osmoregulation overlap overlapping palm patterns petal pigmentation pink predation protection protective reflective rings scale scales sclerites serve shedding size species stone structure thermoregulation types vary