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cape buffalo syncerus caffer african africa wild bovid family bovidae cattle tribe bovini black ruminants grassland savanna mammal swamp floodplains forest water rinderpest plague foot- -mouth disease grass wallows mineral licks kafue national park gregarious lions horns dangerous animal bovines unpredictable temperament big five game trophy hunting robust hooves grazer graze predators kruger herd distress call mobbing behavior fighting dominance nature reserve areas ungulate