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2000 2016 acclimation africa african animal animals antelope arid artiodactyla background became bovidae brown bunch conservation dammah desert ecology enclosures endangered extinct extinction fauna gazelle gemsbok grass grazing group herbivore herd hoofed horn horned horns landscape mammal mammalia natural nature north once oryx ouadi outdoors released rim-ouadi safari sahara sanctuary scimitar species ungulate was white widespread wild wildlife zoo