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brown centipede lithobius forficatus stone common european family lithobiidae arthropod insect invertebrate lithobiid europe chestnut color body segments exoskeleton legs jointed appendages fangs pointed pincers venom prey carnivorous scolopendromorphs upper layers venomous soil rocks rotting logs evasive behavior predator predatory diet spiders slugs worms flies front feeding defence defense mandibles egg molt pair tropical rainforests deserts habitats moist microhabitat water largest terrestrial biomass ecosystems flattened head antennae sharp claws