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alphaketoglutaric acid 2oxoglutaric oxoglutarate alpha ketoglutarate krebs cycle age ageing dietary longevity supplement keto akg ammonia glutarate glutaric ketoglutaric lifespan oxo drawing flat icon hydrogen carbon oxygen organic chemistry formula chemical atomic molecule molecular structure atom compound design vector alpha-ketoglutaric 2-oxoglutaric molecule intermediate metabolite cycle structural model