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euphorbia trigona african milk tree cathedral cactus abyssinian high chaparall euphorbiaceae spurge family flowering plants perennial plant central africa house upright stem green dark three sides thorns pairs two leaves poisonous latex flowers hybrid cuttings root skin irritations pest-free shallow small system ritual hedge herbs tropics shrubs trees succulent cacti convergent evolution indo-malayan region america alternate seldom stipules spines monoecious dioecious milky economic significance