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aculeata animal apocrita arthropod arthropods attack black detail diptera entomology epiponini eusocial fly honey hymenoptera insect invertebrate invertebrates macro myrapetra natural nature nest paper wasp pest plant polistes polistinae pollinate polybia predation prey preying pterygota small spring sting vespidae vespoid vespoidea warrior wasps wild wildlife wing wings adult longwaisted subgenus attacking