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1996 2002 america amphibian animal belize biotope blindness cahuà called cause central cerro common dry eyes family flower food forest foster found freshwater frog grassland habitats hylidae insect insoluble intermittent irritation itzá lake lakes leaf lowland membrane meyer milk moist mucous natural nature noxious observed pain pepper petén produces protected ranging reactions rivers savage secretions severe shore shores sneezing south species sticky subtropical swelling temporary thought trachycephalus tree treefrog tropical veined vermiculatus wildlife