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acacia aepyceros africa allogrooming antelope black black-faced black-tipped bovidae brown browsers bushy coat comb dark deer dental dimorphic diurnal ears ectoparasites etosha eyes faeces family females fruits fur glands glossy grasses grazers gregarious harems herbivorous herds home hornless horns hunting impala leap lyre-shaped melampus melanism midline namibia national nose park petersi pods ranges rare reddish rut savannahs scent self-grooming sexually southwest streaks stripe subspecies tail territorial toothcomb trophy ungulate urine vulnerable water white woodlands