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okapi ungulate mammal endangered species conservation rainforest forest herbivore rare wildlife nature animal african congo zebra giraffe distinctive habitat elusive forest congo basin giraffidae forest-dwelling forest-dwelling conservation efforts wildlife ecosystem biodiversity management protection hoofed elusive canopy undergrowth african biology okapia johnstoni browsing grazer rainforest status iucn okapi reserve ituri understorey basin survival adaptation camouflage poaching hunting natural history habitat loss fragmentation destruction degradation stripes fauna flora sanctuary organizations research education policy behavior reproduction diet canopy undergrowth grazing grazing