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thyroxine t4 thyroid hormone thyroid iodine deiodinase hypothyroidism gland hyperthyroidism thyromimetic thermogenesis hormonal therapy replacement function action endocrine endocrinology level prohormone analogue tyrosine-based metabolism system tissue simple goitre selenium basal metabolic rate protein synthesis carbohydrate protein fat hibernation cycle neuronal activity moulting behaviour body temperature follicular cells thyrotropes anterior pituitary 3d molecule molecular formula skeletal structural structure