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tetrahydroaldosterone steroid lactone mineralocorticoid corticosteroid glucocorticosteroid hormone glucocorticoid receptor inflammation immune system autoimmune disease stress response carbohydrate metabolism protein catabolism anti-inflammatory steroid steroidal endogenous synthetic electrolyte balance fluid mineral collecting tubule intercalated cells hyperaldosteronism hypoaldosteronism congenital adrenal hyperplasia 3d molecule molecular formula skeletal structural structure geometry cell biology biochemistry biomolecule bioactive biochemical biogenesis substrate precursor synthesis derivative bioprecursor