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sulfoquinovosyl distearoylglycerol sulfolipid lipid sulfur-containing functional group sulfoquinovose diacylglycerol sqdg diacylglycerides sulfur cycle sulfatide seminolipid photosynthetic electron transport chain rendering phosphatidylcholine pc phospholipid membrane choline cell arachidonic acid phosphocholine cholesterol fatty ceramide lysophosphatidylcholine bilayer surfactant glycerophospholipid phosphatidylserine lipophilic