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stigmatellin fungicide antifungal fungus antimitotic inhibitor quinol oxidation cytochrome bc1 mitochondria b6f thylakoid membranes complex myxobacterium stigmatella aurantica saccharomyces cerevisiae rhodobacter capsulatus cereibacter sphaeroides paracoccus denitrificans rieske protein metabolite metabolism biosynthesis cycle mechanism pathway path biosynthetic intermediate biogenesis substrate precursor synthesis derivative bioprecursor metabolic enzyme endogenous cell biochemistry biomolecule bioactive biochemical molecule molecular formula skeletal structural structure 3d