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salpichrolid withanolide steroid ergostane secondary metabolite nightshade family tomatillo steroid backbone lactone datura iochroma lycium nicandra physalis salpichroa solanum mandragora withania jaborosa rendering herbal phytochemical phytotherapy plant herb natural herbalism pharmacognosy pharmacopoeia phytoestrogen herbal medicine steroidal endogenous metabolism biosynthesis cycle mechanism pathway path biosynthetic intermediate biogenesis precursor synthesis derivative bioprecursor metabolic