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salicylaldehyde 2-hydroxybenzaldehyde hydroxybenzaldehyde salicylic aldehyde o-hydroxybenzaldehyde 3-hydroxybenzaldehyde 4-hydroxybenzaldehyde isomer chelating agent bitter almond odor buckwheat castoreum castor canadensis fiber beaver perfumery chrysomela populi red poplar leaf beetle metabolite metabolism biosynthesis cycle mechanism pathway path biosynthetic intermediate biogenesis substrate precursor synthesis derivative bioprecursor metabolic enzyme endogenous cell biology biochemistry biomolecule bioactive biochemical molecule molecular formula skeletal structural structure 3d