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s-adenosyl-l-homocysteine sah biosynthetic precursor homocysteine demethylation s-adenosyl-l-methionine adenosylhomocysteinase adenosine amino acid cysteine hyperhomocysteinemia function atherogenesis ischemic injury coronary artery disease 3d metabolite metabolism biosynthesis cycle mechanism pathway path catabolism intermediate biogenesis substrate synthesis derivative bioprecursor metabolic enzyme endogenous contractor human metabolome database hmdb molecule molecular formula skeletal structural structure geometry ball stick atom