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pseudaconitine nepaline extremely toxic alkaloid aconitum ferox ranunculaceae acetylcholinesterase enzyme inhibitor neurotransmitter acraconitine feraconitine pseudoaconitine nepal aconitine veratroylaconine acetylcholine hydrolysis diterpene 3d poison venom toxicant toxicity toxin toxicology poisoning danger hazard acute harmful kill exposure limit antidote intoxication detoxification molecule molecular formula skeletal structural structure geometry ball stick atom