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phosphatidylinositol pi ptdins inositol phospholipid lipid phosphatidylglyceride cell membrane isomer receptor sensory phosphatidylinositol phosphate bisphosphate trisphosphate inositides phosphoinositides amphiphile glycerophospholipid lipid signaling membrane trafficking steric hindrance metabolite metabolism biosynthesis cycle mechanism pathway path catabolism biosynthetic intermediate biogenesis substrate precursor synthesis derivative bioprecursor metabolic enzyme endogenous 3d molecule molecular formula skeletal structural structure biochemical