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phosphatidylethanol peth phospholipid phospholipase pld lipid-gated ion channels alcohol intoxication synthesis marker rendering phosphatidylcholine pc lecithin sphingomyelin phosphatidylethanolamine membrane choline cell arachidonic acid lipid phosphocholine cholesterol fatty ceramide lysophosphatidylcholine bilayer surfactant glycerophospholipid phosphatidylserine lipophilic metabolite metabolism biosynthesis cycle mechanism pathway catabolism biosynthetic intermediate biogenesis substrate precursor derivative bioprecursor metabolic endogenous