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phenylalanine phe essential amino acid c9h11no2 food deficiency group hydroxylase zwitterion dopamine dipeptide breakdown serotonin ionized tyrosine function transamination l-isomer analgesic antidepressant effect precursor dietary phenylketonuria pku boronophenylalanine derivative 3d rendering biochemistry molecule molecular structural structure formula skeletal functional supplement nutrient metabolite metabolism biosynthesis synthesis cycle mechanism pathway aromatic