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nicotinamide adenine dinucleotide nad coenzyme diphosphopyridine nucleotide dpn metabolite metabolism biosynthesis cycle mechanism pathway path catabolism biosynthetic intermediate biogenesis substrate precursor synthesis derivative bioprecursor metabolic enzyme endogenous contractor nucleotide cells nadh redox electrons oxidizing agent reducing agent electron transfer salvage dinucleotide phosphate cofactor anabolic pyrophosphate redox cellular respiration oxidative phosphorylation beta oxidation glycolysis citric acid mitochondrial shuttles mitochondrion transport chain 3d molecule