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malhamensilipin chlorosulfolipids inhibitor protein tyrosine kinase poterioochromonas malhamensis synthesis epoxidation chlorination chrysophyte rendering metabolite metabolism biosynthesis cycle mechanism pathway path catabolism biosynthetic intermediate biogenesis substrate precursor derivative bioprecursor metabolic enzyme endogenous contractor bacteria flora bacterial antimicrobial bactericide pathogen gram-positive gram-negative antibacterial agent microorganism strain molecule molecular structural structure formula skeletal geometry ball stick