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magnesium protoporphyrin mg-protoporphyrin ix porphyrin heme hemoglobin chlorophyll ppix porphine core tetrapyrrole macrocycle aromatic pyrrole ring complexes myoglobin cytochrome porphobilinogen porphyrinogen uroporphyrinogen iii protoporphyrinogen oxidase chelatase heme-like metabolite metabolism biosynthesis mechanism path catabolism biosynthetic intermediate biogenesis substrate precursor synthesis derivative bioprecursor metabolic enzyme endogenous contractor molecule molecular formula skeletal structural structure 3d