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lipoxin a4 lxb4 lx lipoxygenase autacoid metabolite arachidonic acid nonclassic eicosanoid specialized pro-resolving mediators spms polyunsaturated fatty pufa inflammatory response epimer epi-lipoxins neutrophils degranulation biochemistry resolvins neuroprotectins maresins omega acids eicosapentaenoic docosahexaenoic transcellular biosynthetic pathway supplement prostaglandin endocannabinoid system rendering metabolism biosynthesis path catabolism biosynthetic intermediate biogenesis substrate precursor synthesis derivative bioprecursor metabolic enzyme endogenous