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lathosterol cholesterol-like molecule sterol-c5-desaturase lathosterolosis cholesterol biosynthesis metabolic diseases dyslipidemia lipoprotein intermediate rendering metabolite metabolism cycle mechanism pathway path catabolism derivative bioprecursor metabolic enzyme endogenous contractor phosphatidylcholine pc lecithin sphingomyelin phosphatidylethanolamine phospholipid membrane choline cell arachidonic acid lipid phosphocholine cholesterol fatty ceramide lysophosphatidylcholine bilayer surfactant glycerophospholipid phosphatidylserine lipophilic