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lanostane polycyclic hydrocarbon trimethylcholestane triterpene monoterpenoid terpenoid monoterpene terpene cucurbitane tautomer isomer stereoisomer steroid biogenetic precursor 3d essential oils biosynthetic skeletal isoflavone path ball stick herbalism dietary prescription drug cell molecular biological activity flavone coumarin biosynthesis flavonoid mechanism cycle biochemistry intermediate biomolecule pharmacognosy herbal formula healthy aging phytochemical traditional chinese medicine bioactive natural antioxidant alkaloid metabolite